Tyrannosaurus rex was one of the largest terrestrial predators that ever existed. The species lived during the final segment of the Mesozoic Era, known as the Late Cretaceous, specifically the late Maastrichtian age. This timeframe, spanning approximately 69 to 66 million years ago, placed the massive carnivore at the close of the Age of Dinosaurs. Its existence was confined to a North American landscape dramatically different from the continent we know today.
The Geography of Laramidia
The geographic domain of T. rex was an ancient, isolated landmass called Laramidia, which formed the western portion of what is now North America. This region existed as an island continent, a long, narrow strip of land stretching from present-day Alaska down to Mexico. Laramidia was defined by the presence of a vast, shallow body of water known as the Western Interior Seaway (WIS).
The Western Interior Seaway bisected the continent, separating Laramidia from a smaller, eastern landmass called Appalachia. This marine barrier dictated the eastern boundary of the T. rex range. Geological evidence for this environment is concentrated in rock layers like the Hell Creek Formation and the Lance Formation, spanning parts of Montana, North Dakota, and Wyoming. These fossil-rich formations represent coastal plain sediments deposited along the retreating edge of the seaway, providing a detailed record of the dinosaur’s homeland.
Climate and Water Systems
Laramidia was characterized by a warm, humid, and subtropical climate, contrasting sharply with modern North America. Global temperatures were significantly higher during the Late Cretaceous, creating a greenhouse world. Mean annual temperatures were estimated to be between 16 and 20 degrees Celsius in the northern latitudes. This thermal stability meant the region experienced minimal seasonal temperature fluctuation.
The landscape’s hydrology was dominated by extensive coastal plains and low-lying areas frequently inundated with water. The environment consisted of vast floodplains, large meandering rivers, marshes, and coastal swamps that drained into the Western Interior Seaway. Fossil evidence, such as the abundance of ancient crocodile relatives, supports this perpetually wet, low-elevation environment. Sediment flow from the newly rising Rocky Mountains fed these river systems, creating the swampy, deltaic conditions that preserved the rich fossil record.
The Late Cretaceous Ecosystem
The lush, waterlogged landscape supported a diverse biological community. The flora was highly developed, with flowering plants (angiosperms) making up roughly 90 percent of the vegetation. Dense forests and open woodlands included conifers, ferns, and cycadophytes. This abundance of angiosperms provided a rich food source for the large herbivorous dinosaurs that shared the environment with T. rex.
The fauna of Laramidia featured large plant-eaters that served as prey for the apex predator. Among the most recognizable were the horned dinosaur Triceratops and the duck-billed dinosaur Edmontosaurus, both common in the northern part of the continent. Heavily armored herbivores, such as Ankylosaurus, also inhabited the region, presenting a challenging target with bony plates and tail clubs.
The ecosystem also included numerous smaller species that completed the Late Cretaceous food web.
Smaller Fauna
Small, feathered theropods like dromaeosaurids
Pachycephalosaurs with dome-shaped skulls
Various early species of mammals, most smaller than a rabbit
The rivers and swamps were populated by turtles, lizards, and crocodilians, while the skies were patrolled by pterosaurs. This complex web of life defined the world in which Tyrannosaurus rex reigned as the ultimate predator.

